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Claire Holt portrays original vampire Rebekah Mikaelson.
Main article: Original Vampires (The Vampire Diaries)
Rebekah Mikaelson is Klaus and Elijah's younger sister and also an original vampire. Rebekah shares a deep bond with her brother Klaus, and was the only family member that remained loyal to him despite his vicious behavior. Klaus thus kept her by his side, while he kept the rest of his family staked and stored away in boxes for hundreds of years. She had a romantic interest in Stefan Salvatore during their encounters in the early 1920s. However, when Klaus and Rebekah were once again forced to go on the run after the arrival of their vengeful father Mikael, Rebekah professed an interest to stop going on the run and live a stable life with Stefan by her side. Klaus subsequently forced her to choose between him and Stefan, and she chooses the latter. Klaus then daggers Rebekah.
He brings her back to life during the period in which the show takes place, in order to obtain her necklace (which is originally Esther's). The same necklace had been gifted to Elena by Stefan in season one. Rebekah and Klaus thus head to Mystic Falls. Rebekah develops a deep hatred and envy of Elena, and continually tries to hurt, kill, and emotionally distress her, until her brother makes her stop. Later on, Rebekah is once again daggered by Elena, to simplify the Salvatores' plans of killing Klaus. Elena makes a deal with Klaus for her to reveal Rebekah's location, while Klaus promises not to harm Jeremy. He takes her body with him. She is eventually undaggered by Elijah. During her time in Mystic Falls, Rebekah enrolls at the local high school, much to the disgust of Elena and her friends. There, she at first makes romantic advances towards Tyler Lockwood, Caroline Forbes's boyfriend. She later develops a fondness for Matt Donovan, and invites him to her family's ball. Rebekah is shown to be lonely and insecure, though she hides this by being cruel and violent. This is demonstrated quite well through her sexual rendezvouses with Damon Salvatore, an enemy of her family, and the subsequent torture she makes him endure upon being betrayed by him. Rebekah's body is temporarily taken over by her witch mother, Esther, through a spell which Esther performs. She invades Rebekah's body in order to be able to go unnoticed while scheming to kill all of her Original children. Esther later leaves Rebekah's body and stakes her. However, Klaus restores Rebekah to full vitality by removing the stake. In the season 3 finale, while Matt is driving Elena back to Mystic Falls, after trying to escape with her, Rebekah is seen standing in the middle of Wickery Bridge. Matt sees her, loses control, and drives the car in the lake, which ultimately results in Elena's death and subsequent transformation into a vampire.
In the season 4 premiere, Rebekah is captured alongside every other vampire in Mystic Falls. While en route to their detainment, Caroline is rescued by Klaus, inhabiting Tyler's body, but leaves his sister there. While locked up, Rebekah realizes that Elena is in transition but hasn't fed on human blood, which she is delighted about because "I get to watch you die all over again". However, as she sits in her cell, she listens to Elena and Stefan's heartfelt confessions of regret and love to each other with a conflicted look on her face. Rebekah then helps Stefan kill one of their guards, allowing Elena to feed on his blood and complete the transition into a vampire. After being released, Rebekah storms home and confronts Klaus, now back in his original body. Rebekah, in tears and furious, tells him how she always loved him despite everything he has done but he doesn't love or care for her as a sister. Rebekah then destroys the last few bags of Elena's blood, preventing him from creating more hybrids. In retaliation, Klaus disowns Rebekah, telling her he does not love her and she is not his sister anymore, and breaks her neck. Rebekah leaves Klaus' mansion and moves into her own house. She befriends April Young, an orphan who moves back to Mystic Falls after her father is killed in an explosion. She is then asked by Stefan to tell him and Klaus the story of the Five and the cure for vampirism. After Klaus and Stefan find out what they need from her, Klaus daggers Rebekah and has Stefan hide her.
Later, April learns from Caroline where Rebekah is and un-daggers her. With April's help, Rebekah takes Elena, Stefan and Caroline prisoner so that she can learn what progress they have made in finding the cure. After toying with them and finding out what she wants, she releases them. Stefan later approaches her where she states she still wishes to find the cure so she can use it on Klaus. Stefan then proposes the two work together to get the cure since they are the only two without an edge, since Klaus has the sword, Damon is with Jeremy, and Bonnie is working with Professor Shane. In the episode "A View to Kill" Rebekah and Stefan work together to find the headstone of the first immortal creature. She then convinces him to stop caring about Elena and the two have sex. Throughout season 4 Rebekah is shown to be interested in Matt Donovan but he is not interested in her. Near the end of the season the four spend more time together and appear to be friends. Whilst Rebekah is on a test to see if she could be human Matt tells her that she wouldn't be a good human because she's not a good person. Later on Rebekah saves Aprils life which changes Matt's view of her. In the season 4 finale, Rebekah and Matt kiss while Matt is being held hostage and Rebekah saves his life, taking his place and allowing him to leave in order to attend graduation. Matt then develops some feelings for Rebekah and agrees to spend the summer traveling the world with her, saying that whatever happens on the road stays on the road.
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